BTT Morfologi Vegetatif

download BTT Morfologi Vegetatif

of 26

Transcript of BTT Morfologi Vegetatif

  • 8/10/2019 BTT Morfologi Vegetatif

    1/26

    2

    GBI 353

    Botani Tumbuhan Tinggi (3 SKS)Drs. Endang Dayat, M.Si.Drs. Didi Jaya Santri, M.Si.

    Program Studi Pendidikan Biologi

    Fakultas Keguruan dan Ilmu Pendidikan

    Universitas Sriwijaya

    2005

  • 8/10/2019 BTT Morfologi Vegetatif

    2/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Meristem Batang

    Meristemmerupakan pusat pembelahan

    sel mitosis yang menghasilkan jaringan

    tumbuhan

    Meristem apicalatau terminalmenghasilkan jaringan akardan taruk

    yang merupakan dua sistem organ pada

    tumbuhan berpembuluh

    Structures dan terminologi botani baik

    vegetatif maupun reproduktifberhubungan dengan produk meristem

    taruk apikal, dan pola aktivitasnya

    menentukan kerangka dasar organisasi

    struktural tumbuhan (see OSU Botany

    300, for more detail).

    Batang

    http://www.hcs.ohio-state.edu/hcs300/cell2.htmhttp://www.hcs.ohio-state.edu/hcs300/cell2.htmhttp://www.hcs.ohio-state.edu/hcs300/cell2.htmhttp://www.hcs.ohio-state.edu/hcs300/cell2.htm
  • 8/10/2019 BTT Morfologi Vegetatif

    3/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    As indicated by both size and longevity of some vascular plants, theapical meristem is indeterminate. Its activity of producing cells, orlaying the bricks of the plant body, can continue indefinitely.

    The apical shoot meristem also has the capability to producesecondary centers of meristematic activity, lateral meristems. The

    lateral meristems provide a structural groundplan for mapping theangiosperm shoot.

    The position of lateral or axillary meristems, as either lateral buds orsecondary shoots and often subtended (positioned beneath) by aleaf, is known as a node.

    Shoot tissue that extends between these points is knowned as theinternode

    Batang

  • 8/10/2019 BTT Morfologi Vegetatif

    4/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang Aside from the lateral meristem, another

    secondary meristematic product of theshoot meristem - the vascular cambium- is significant. This is a column ofdividing cells that produces vasculartissue (xylem and phloem).

    Since meristematic activity of the

    vascular cambiumvaries according toavailable resources and conditions duringthe growing season, its product - wood -often takes the form of 'annual rings'.

    Angiosperms with an active vascularcambium are 'woody' and those taking atree-like form (single stem greater than 4-5 m in height) are arborescentwhereaswoody angiosperms with a shrub-likeaspect (more than 1 shoot, less than 4-5m) are fruticose.

    Most angiosperms lack a vascularcambium, i.e., they are non-woody herbs

    or herbaceous.

  • 8/10/2019 BTT Morfologi Vegetatif

    5/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

  • 8/10/2019 BTT Morfologi Vegetatif

    6/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang

    Both woody and herbaceous angiosperms can showindeterminate apical meristem activity and a perenniallifestyle. Herbaceous perennials living in temperate or seasonableparts of the world sacrifice 'above ground' tissue during thedifficult season but often retain dormant meristems andphotosynthate in a tough, underground structure known as a

    caudex. However, most herbaceous flowering plants are annuals in that

    they show an annual cycle of seed germination, vegetativegrowth, and reproduction with the next generation passing the'difficult' annual period (Winter, dry season) within the seed orfruit.

    Some angiosperms dedicate a full growing season to vegetativegrowth, sequester photosynthate - often in undergroundstructures - through the difficult season and invest this in areproductive effort at the start of the second year. Thesebiennialsare often found at the market (carrot, onion, etc.)because we harvest the saved resource prior to its 'intended'use by the plant.

    Life Style

  • 8/10/2019 BTT Morfologi Vegetatif

    7/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang

    Many herbaceous angiosperm genera of central Texas includespecies with varying 'life styles' (annual, binnual, perennial) and thiscan be a factor or 'key character' for identification.

    Thus, it is important to insure that the sample collected includes a'below ground' component.

    Also, terms are discrete and biological variation can be continuousand, as a result, difficult to define using specificterms. Consequently, 'bridging' terms can be applied.

    The term suffrutescent, for instance, denotes an herbaceousperennialthat shows some woodytendencies, usually at the base.

  • 8/10/2019 BTT Morfologi Vegetatif

    8/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Pohon Herba daun lebar

    Herba daun sempitPerdu

    Sukulen

  • 8/10/2019 BTT Morfologi Vegetatif

    9/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang

    The stem 'caul-' refers to the primary product of the terminalmeristem, the shoot. Caulescentdenotes a 'normal' shoot withnodes and discernable internodes. Acaulescent, on the other hand,descibes a situation where the internodes are reduced and, as aresult, this shoot is compressed to the point that the plant looks'stemless', such as head lettuce (Lactuca- Asteraceae) and

    cabbage (Brassica- Brassicaceae). Terms relating to variousadaptive permutations of the shoot include: Erect- 'normal' growth toward the sun

    Prostrate- a horizontal shoot, flush to the ground

    Decumbent- a horizontal shoot, but not flush to the ground - the tipspoint upward

    Caespitose- producing clustered, multiple shoots forming tufts orcushions

    Scandent- twining, climbing - vine-like

    Scapose- acaulescent but producing an erect, leafless flowering stalk(scape) - Taraxacum off ic inale

    http://caliban.mpiz-koeln.mpg.de/~stueber/lindman/36.jpghttp://caliban.mpiz-koeln.mpg.de/~stueber/lindman/36.jpg
  • 8/10/2019 BTT Morfologi Vegetatif

    10/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang

    Rhizome- horizontal, elongate andunderground, often thick and fleshy (herbaceousperennial are sometimes rhizomatoussee:Cyperus- Cyperaceae)

    Corm- vertical, compressed and underground,often thick and fleshy (herbaceous perennial)

    Bulb- acaulescent with expanded, fleshy leafbases (Onion - Al l ium- Liliaceae)

    Function modifications of the shoot include:

    http://www.csdl.tamu.edu/FLORA/tfplab/veg11.jpghttp://www.csdl.tamu.edu/FLORA/fa08/fa08039.jpghttp://www.csdl.tamu.edu/FLORA/fa08/fa08039.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/veg11.jpg
  • 8/10/2019 BTT Morfologi Vegetatif

    11/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang

    Tuber- thick and fleshy and usuallyunderground, like a rhizome, butcompressed - via reduction of the internodes- and enlarged as a water/photosynthate

    storage structure (Potato - Solanum-Solanaceae)

    Stolon- horizontal, above-ground, oftenrooting at the nodes - strawberry (Fragaria-Rosaceae) and St. Augustine grass

    (Stenotaphrum

    - Poaceae) arestoloniferous.

    Thorn- a hard, sharp-pointed modified shoot(Honey locust - Gleditsia-Caesalpiniaceae)

    http://www.csdl.tamu.edu/FLORA/tfplab/veg13.jpghttp://csdl.tamu.edu/FLORA/tfplab/veg12.jpghttp://www.csdl.tamu.edu/FLORA/fa03/fa03067.jpghttp://www.csdl.tamu.edu/FLORA/fa03/fa03067.jpghttp://csdl.tamu.edu/FLORA/tfplab/veg12.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/veg13.jpg
  • 8/10/2019 BTT Morfologi Vegetatif

    12/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Batang

    Another feature of the shoot that can beused as a key character is the nature ofthe central tissue or pith.

    The pith is normally present as soft,

    uniform tissue (continuous), but thiscan be intersected by discs of moredense tissue (diaphragmed) with, insome cases, the 'pithy' soft tissueabsent (chambered).

  • 8/10/2019 BTT Morfologi Vegetatif

    13/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Daun Their structure and position on the stem also provides a rich suite of

    taxonomic characters. The fossil record suggests that the first angiosperms had simple,

    alternate, entireleaves.

    STRUCTURE: (overview) Simple- blade a single, undivided structure - example

    Compound- blade divided into substructures (leaflet), each with

    its own stalk (petiolule)

    Pinnately Compound- leaflets arranged along a main axis (rachis) -

    example

    'Even' = no terminal leaflet,

    'Odd' with a terminal leaflet

    Bipinnately Compound- as above but leaflets compounded -

    example

    Palmately Compound- leaflets with petiolules attached to a single

    point - example

    http://www.csdl.tamu.edu/FLORA/Wilson/tfp/veg/leaftypess01.htmhttp://www.csdl.tamu.edu/FLORA/tfplab/lfpart.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/1pinn.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/2pinn.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/palm.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/palm.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/2pinn.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/1pinn.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/lfpart.jpghttp://www.csdl.tamu.edu/FLORA/Wilson/tfp/veg/leaftypess01.htm
  • 8/10/2019 BTT Morfologi Vegetatif

    14/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Daun

    Alternate- a single leaf at the node

    Opposite- two leaves at the node, each usually subtending

    (immediately below and close to) a lateral bud or shoot

    Whorled orverticillate - more than two leaves at the node

    Rank - when applied to leaves, denotes rows of leaves along the

    shoot

    Decussate- four ranked or opposite leaves alternating at right

    angles to those above and below Cauline - leaves are those associated with the central shoot

    Fascicled- closely clustered or grouped

    PHYLLOTAXY (overview)

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    15/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Daun

    Structural features of a simple leaf, as depictedabove include the stalk (petiole) which is usuallyattached to the shoot at the node, subtending(attached beneath) the lateral meristem, andoften with a distinctive basal enlargement

    (pulvinus). Foliar or leaf-like appendages that are not similar

    to the typical photosynthetic leaf, but occur onthe plant, often in association with flowers orinflorescences, are known as bracts.

    When they are associated with the petiole base,

    often flanking it at the node, they are calledstipules.

    Significant indentations in the leaf blade tissueare called sinuses.

  • 8/10/2019 BTT Morfologi Vegetatif

    16/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Daun

    Linear- long and narrow with nearly parallel sides

    Lanceolate- Lance-shaped, tapering from a broad base to an apex;

    much longer than wide Oblong- long and wide with nearly parallel sides

    Elliptical- football (American) shaped - a flatten circle, usually morethan twice as long as wide

    Ovate- egg-shaped with the broadest part toward the base (notethat obovate is the reverse relative to the point of petiole attachmentto the blade)

    Cordate- heart shaped with a basal sinus

    LEAF BLADE SHAPES General (overview)

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    17/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Daun

    Sagittate- basal lobes pointed back toward the petiole- arrowhead shaped

    Auriculate- with ear-lobe like appendages at the base Petiolate- with a petiole (three of the overview images)

    Sessile- petiole reduced - the leaf blade appears to beattached to the stem

    Hastate- basal lobes pointing outward relative to thelong axis of the blade

    Oblique dasar daun tidak simetris bilateral

    LEAF BLADE SHAPES Basal Lobing (overview)

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    18/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Deltoid- triangular

    Peltate- shield shaped with the petiole not attached at the

    blade margin Reniform- kidney shaped

    Daun

    LEAF BLADE SHAPES panjang x lebar (overview)

    Palmate Pinnate

    Divided (also 'parted' (almost divided) and

    'pinnatifid' (pinnately parted)

    LEAF BLADE lobing (overview)

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    19/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Dichotomous- equal bifurcation or forking, more common in

    ferns and gymnosperms

    Palmate- the main veins radiating from the point of petiole/bladejunction

    Pinnate- a central 'mid-vein' or -mid-rib' with with lateral veins

    arising along its length

    Parallel- veins running side by side without intersecting - typical

    of monocots Reticulate- typical pattern of dicots with intersecting network of

    veins

    LEAF BLADE Venation (overview)

    M f l i V if

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    20/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Acuminate- tapering gradually to a prolonged point

    Acute- ending at a point that is less than a right angle -distinct/sharp (not acuminate)

    Obtuse - with a blunt or rounded tip Cuspidate- tipped with a sharp and rigid point

    Mucronate- tipped with a small, pointed extension of the mid-rib

    Truncate- cut squarely across the apex

    Retuse- having a shallow notch at the apex Emarginate- also notched at the apex, mid-way between

    Retuse and Cordate

    Cordate- a deep notch at the apex, found in an obcordate leaf

    LEAF BLADE Tips (overview)

    M f l i V t tif

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    21/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Entire- no indentations, lobes, or teeth - smooth

    Serrulate- small, marginal teeth pointing toward the blade apex

    Serrate- marginal teeth pointing toward the blade apex (saw-like) Doubly serrate- with both small and larger serrations

    Dentate- marginal teeth point outward

    Spinose-dentate- as above with the teeth point-tiped

    Crenate- with low rounded or blunt teeth

    Crenulate- with small, low rounded or blunt teeth Undulate- a slightly wavy margin

    Broadly crenate - intermediate between undulate and sinuate

    Sinuate- deeply wavy margin

    LEAF BLADE Margins (overview)

    M f l i V t tif

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    22/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Puberulent- minutely pubescent (covered with short, softtrichomes

    Stellate- with star-shaped trichomes

    Tomentose- woolly in appearance, densely matted, softtrichomes

    Glandular- with glands (trichomes with secretory function )

    Hirsute- long shaggy trichomes, often stiff or bristly to the touch

    Surface Feature - Trichomes (overview)

    M f l i V t tif

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    23/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Scabrous- thin, membranous, dry and often rough

    Strigose- stiff hairs appressed and all pointing in one direction

    Villous- dense covering of long, fine, soft trichomes Pilose- scattered covering of long, fine, soft trichomes

    Glabrous- no trichomes - smooth surface

    Surface Feature - Trichomes (overview)

    M f l i V t tif

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    24/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Glaucous- have a waxy bloom(photo)

    Coriaceous- a rough, leathery texture

    Tap- the central or primary root that enlarges and grows downward

    - typical of dicots (photo)

    Fibrous- thread like, often tough root system - typical of monocots

    (photo)

    Adventitious- developing from something other than the root

    meristem or radical(photo)

    Surface Feature - Other (overview)

    Root Meristem (overview)

    M f l i V t tif

    http://www.csdl.tamu.edu/FLORA/tfp/prunfrt3.jpghttp://www.csdl.tamu.edu/FLORA/hdwimages2/hdwt4-57.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/veg2.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/veg5.jpghttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/tfplab/veg5.jpghttp://www.csdl.tamu.edu/FLORA/tfplab/veg2.jpghttp://www.csdl.tamu.edu/FLORA/hdwimages2/hdwt4-57.jpghttp://www.csdl.tamu.edu/FLORA/tfp/prunfrt3.jpg
  • 8/10/2019 BTT Morfologi Vegetatif

    25/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif

    Pedicel- stalk of an individual flower if part of an inflorescence(cluster of flowers)

    Peduncle- stalk of an inflorescence or that of a solitary flower

    Calyx - lower-most whorl of floral appendages (made up ofsepals)

    Corolla- next flora whorl or series of appendages up from thecalyx (made up of petals)

    Perianth- term applied to the non-reproductive floralappendages

    Androecium- above the corolla and the first reproductivewhorl (composed of stamens)

    Androperianth - terms applied to perianth and androeciumcombined

    Gynoecium- last of the series of appendages and positionedabove the others (composed of carpels)

    Reproductive Structure (overview)

    Morfologi Vegetatif

    http://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.htmlhttp://www.csdl.tamu.edu/FLORA/201Manhart/veg/leaf.arrange/leafarrange.html
  • 8/10/2019 BTT Morfologi Vegetatif

    26/26

    otani Tumbuhan Tinggi

    Morfologi Vegetatif